Former champ Matlala, who has a BComm degree, joined Milenkovic's Branco Sports Productions (BSP) in early 2005 as a co-director, giving his name in return for a 25% stake.
But electronic company records show that BSP has just one director, Susanna Susara Maree.
"I think that's Branco's wife," said Matlala, a winner of four world titles at flyweight and junior-flyweight.
He claimed he had never received bonus or dividend payments, yet he had helped secure hefty sponsorships, especially from various provincial governments.
"We would raise between R2-million and R4-million for a show, and I would get a cut of R50000 or R60000. For tournaments where we didn't get government sponsorships, I'd get R3000. For motivational talks I get more than that, and sometimes that's just for one hour."
Milenkovic said Matlala had yet to tell him that he had quit.
"We spoke on the phone on Wednesday night and there was no expression of unhappiness. I've tried to call him since and he's not taking my calls. To say I'm disappointed would be an understatement.
"If I gave the impression that you could become a millionaire overnight in boxing, then I'm sorry. I'm still paying for my house."
Milenkovic said he could not comment further on Matlala's allegations.
However, the promoter said he believed there were other reasons for Matlala's departure. "I've learnt that behind this is outside interference. I have the names of two promoters who are influencing him and I will release those names at a later date. I hope Jake is not being used. Obviously there is motivation behind his actions. I cannot comment further."
Matlala said his wife had recently taken out a promoter's licence and he would soon join her in the new venture, Baby Jake's Baby Promotions.
He added that his lawyers were looking at the BSP issue.
Meanwhile, this week Milenkovic received a lawyer's letter from boxer Daniel Bruwer, who refused to fight on Milenkovic's bill on Friday night.
Bruwer's camp claimed that his contract for the fight was invalid for various reasons, including that the number of rounds had been altered from 12 to 10.
Milenkovic admitted he made the change, but said the boxer had agreed to it when it appeared the fight would lose its title status. But his R65000 purse would have remained the same.
He said Bruwer had come to him "with tears in his eyes" because he'd last fought in August 2008, on a bill promoted by Rodney Berman. He then joined promoter King Ramathe, but failed to get a fight.
"There are people trying to manipulate other people to get to me because of my success," said Milenkovic. "This is boxing and I've just got to roll with the punches."
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